Cynodonichthys     (CAS - Catalog of Fishes)

  

Untergattung Meek 1904 / Gattung Costa 2011 / Untergattung Huber 2012

 

Type species Cynodonichthys tenuis (Meek, 1904)

 

Vomerivulus (subgenus of Rivulus) Synonym of Cynodonichthys Meek 1904

Phylogenetic position and taxonomic status of Anablepsoides, Atlantirivulus, Cynodonichthys, Laimosemion and Melanorivulus (Cyprinodontiformes: Rivulidae)

Wilson J. E. M. Costa 2011

A maximum parsimony analysis of a combined set of mitochondrial and morphological data available in the literature for 33 rivuline taxa and three outgroups confirms Rivulus as a paraphyletic assemblage. In order to adjust a generic classification to our present phylogenetic knowledge on rivuline relationships, the following taxonomic changes are proposed: Rivulus, restricted to two species endemic to Cuba, is hypothesized to be the most basal rivuline lineage, distinguished from all other non-annual rivulines by having all hypurals fused into a single plate, neural prezygapophysis of caudal vertebrae rudimentary, fourth ceratobranchial teeth absent, about 50 % of the anterior portion of the caudal fin covered by scales, four neuromasts on the anterior supraorbital series, and a black round spot with white margin on the dorsoposterior portion of the caudal peduncle in females; Anablepsoides, Atlantirivulus, Laimosemion, Melanorivulus and Cynodonichthys, previously classified as subgenera of Rivulus, are considered as valid genera; Laimosemion, including 24 species from northern South America, constitutes the sister group to a clade comprising Melanorivulus, Cynodonichthys, Anablepsoides, Atlantirivulus, and all annual rivuline genera, which is supported by a well-developed dorsal process of the urohyal and an expanded lateral articular facet of the first hypobranchial; Melanorivulus, comprising 34 species from central and northeastern South America, Cynodonichthys with 27 species from Central America and Trans-Andean South America, Anablepsoides with 42 species from northern and northeastern South America and Smaller Antilles, and Atlantirivulus, with 11 species endemic to the eastern Brazilian coastal plains are diagnosed by combinations of morphological characters, including osteology, cephalic laterosensory system and colour patterns.

Huber, J.H., Reappraisal of the Phylogeny of Rivulus and its Allied focused on External Characters. Killi‐ Data Series 2012, 9‐25, 3 figs., 2 tabs.

Abstract: In the present study all known to date phylogenetic lineages encompassing the genus Rivulus in the large sense and its relatives, 44 in total, are benchmarked against a very long list of external characters involving global and detailed morphology of each, plus some characters related to biology, behavior and ecology, plus a few characters concerning the vertebrae counts and their structure. The parsimony analysis for these 144 characters results in 3 trees only, very much in line with the global stability of major lineages and the consensus tree produces bootstrap values that are much higher than previously released studies with morphological and osteological characters. In the resulting tree, outside Kryptolebias already separated in 2004, the genus Rivulus in the large sense is shown as a monophyletic assemblage like in the molecular study by Murphy, Thomerson & Collier (1999), with annual and non‐ annual species separated, but unlike Costa (2011) who used both morpho‐osteological and molecular combined characters. Apart from the coherence of the genus Rivulus in a large sense, the analysis newly suggests the primitive position of Millerichthys, the monophyly of major annual subclades, and the phylogenetic relationship of a clade of 3 subgenera of Rivulus, namely Prorivulus, Atlantirivulus, Melanorivulus in line with a previous palaeo‐bridge hypothesis and the minor osteological diagnostic characters that were forwarded for these 3 taxa should be deepened and re‐evaluated. Two other subunits of Rivulus, namely Anablepsoides and Oditichthys, are atypically placed in the tree, but their 2 type species were not studied by Costa (2011) and that too would need re‐evaluation. Finally several phylogenetic lineages belonging to the huge ‐and very poorly known‐ Amazon basin ends up in an unstable position and more material and characters are thought necessary for further research of this very difficult group of fish.

 

 

Cynodonichthys Arten (32 Arten)

 

Cynodonichthys azurescens (Vermeulen, 2013) 

Four new species of Rivulus Poey, 1860, subgenus Cynodonichthys Meek, 1904 (Teleostei: Cyprinodontiformes: Rivulidae) from the Magdalena River Basin, Central Colombia, including notes on Rivulus (Cynodonichthys) elegans Steindachner, 1880. Killi-Data Series, 2013: 42-63.

 

Cynodonichthys birkhahni (Berkenkamp & Etzel, 1992)

Die Rivulus-Arten von Panama. 2. Wiederbeschreibung von R. brunneus & Beschreibung von R. birkhahni spec. nov. von Chiquiri Grande. D.K.G. (Deutsche Killifisch Gem.) J., 24 (7): 102, figs.

Type locality: near Chiriqui Grande, provincia Bocas del Toro, Panama, Caribbean side.

 

Cynodonichthys boehlkei (Huber & Fels, 1985)

Costa, W.J.E.M., 2003. Rivulidae (South American Annual Fishes). p. 526-548. In R.E. Reis, S.O. Kullander and C.J. Ferraris, Jr. (eds.) Checklist of the Freshwater Fishes of South and Central America. Porto Alegre: EDIPUCRS, Brasil. 

South America: Magdalena River basin, Colombia.

 

Cynodonichthys brunneus (Meek & Hildebrand, 1913)

New Species of Fishes from Panama. Field Columbian Mus. Zool. Ser. (Chicago Nat. Hist. Mus. Fieldiana Zool.) Publication, 166, 10 (8): 86.

 

Type locality: Toro Point (Upper Reservoir creek), Canal Zone, (Atlantic) Panama.

 

Cynodonichthys chucunaque (Breder, 1925)

New Loricariate, Characin, and Poeciliid Fishes from the Rio Chucunaque, Panama. Amer. Mus. Novitates, 180: 7, fig. 6.

 

Type locality: río Chucunaque, in a small side stream near Yavisa, Darien Province, Panama.

Cynodonichthys chucunaque "PAN 2012-01"

 

Cynodonichthys degreefi (Collier, 2016) 

The Rivulus of Guatemala: Rivulus tenuis Meek, 1904, Rivulus godmani Regan, 1907 and Rivulus degreefi sp. nov. Journal of the American Killifish Association, 49 (2): 34-47.

 

Cynodonichthys elegans (Steindachner, 1880)

Zur Fischfauna des Cauca und der Flüsse bei Guayaquil. Denk. Akad. Wiss. Wien, 42: 85, pl. 6 (fig. 6).

Type locality: río Cauca (near Caceres, subseq.), northern Colombia.

 

Cynodonichthys frommi (Berkenkamp & Etzel, 1993)

Die Rivulus-Arten von Panama. 4. Rivulus frommi spec. nov., eine neue Bachlingsart von El Valle, Zentral Panama. D.K.G. (Deutsche Killifisch Gem.) J., 25 (6): 81, figs.

Type locality: Coclé Province, north of El Valle, central Panama.

 

Cynodonichthys fuscolineatus (W. A. Bussing, 1980)

Status of the Cyprinodontid Fish Genus -Rivulus in Costa Rica with Descriptions of new endemic Species. Brenesia, San Jose, 17: 336, figs. 1A, 5A, 6A.

 

Type locality: quebrada (creek) Santa Fe, 6.1 km north of Tilaran on road to Aguacate, Guanacaste Province, (northern) Costa Rica, 640 m altitude.

 

Cynodonichthys fuscolineatus "Senderos Oasis Cano Negro CR"

 

Cynodonichthys glaucus (W. A. Bussing, 1980)

Status of the Cyprinodontid Fish Genus -Rivulus in Costa Rica with Descriptions of new endemic Species. Brenesia, San Jose, 17: 342, figs. 2D, 5D, 6D.

Type locality: tributary of río Pedregoso at southern edge of San Isidro de El General, San José Province, (western Pacific) Costa Rica, 680 m. altitude.

 

Cynodonichthys godmani (Regan, 1907) 

Pisces. In: Biologia Centrali America. P. Press, 8: 65.

Type locality: Guatemala (without details ; lower rio Motagua or El Peten, subseq.).

Collier, G.E. (2016): The Rivulus of Guatemala: Rivulus tenuis Meek, 1904, Rivulus godmani Regan, 1907 and Rivulus degreefi sp. nov. Journal of the American Killifish Association, 49 (2): 34-47

 

Cynodonichthys hendrichsi (Álvarez & Carranza, 1952)

Cuatro Especies Nuevas de Peces Dulceacuicoles del Sureste Mexico. Ciencia, 11 (10-12): 281, figs. 1-2.

Type locality: 2 km southeast of Salto de Agua, Chiapas (about 80 km southeast Villahermosa), southeastern Mexico.

 

Cynodonichthys hildebrandi (G. S. Myers, 1927)

A new Catalog of the Freshwater Fishes of Panama. Field Columbian Mus. Zool. Ser. (Chicago Nat. Hist. Mus. Fieldiana Zool.) Publication, 22: 316, fig. 8.

Type locality: laguna Grande (Davis, today David) near Volcán, Río Chiriquí Viejo basin, western Panama, 1500 m altitude.

 

Cynodonichthys isthmensis (Garman, 1895)

The Cyprinodonts. Mem. Mus. Comp. Zool. Harvard, 19 (1): 140.

Type locality: río San José, Costa Rica.

 

Cynodonichthys kuelpmanni (Berkenkamp & Etzel, 1993)

Die Rivulus-Arten von Panama. 3. Rivulus kuelpmanni spec. nov., eine neue Bachlingsart von Punta Pena. D.K.G. (Deutsche Killifisch Gem.) J., 25 (2): 17, figs.

Type locality: at foot of Cordillera Central, 20 km from Punta Peña, provincia Bocas del Toro, Atlantic Panama, 200 m altitude.

 

Cynodonichthys leucurus (Fowler, 1944)

Freshwater Fishes from Northwestern Colombia. Proc. Acad. Nat. Sci. Philad., 96: 244, figs. 21-23.

Type locality: upper río Juradó (about 40 km north of Jurado city, on the Pacific coast, subseq.), northwestern Colombia, 200 m altitude.

 

Cynodonichthys magdalenae (C. H. Eigenmann & Henn, 1916)

On various South American Poeciliid Fishes. Ann. Carnegie Mus., 10 (9): 109.

Type locality: Ibagué (near upper rio Magdalena), Colombia, 1300 m altitude.

 

Cynodonichthys monikae (Berkenkamp & Etzel, 1995)

Die Rivulus-Arten von Panama. 5. Rivulus monikae spec. nov., eine weitere neue Bachlingsart aus den bewaldeten und bergigen Teilen der Provinz Cochlé, Zentral Panama. D.K.G. (Deutsche Killifisch Gem.) J., 27 (1): 1, figs.

Type locality: 15 km north of La Pintada, Coclé province, central Panama.

 

Cynodonichthys montium (Hildebrand, 1938)

A new Catalog of the Freshwater Fishes of Panama. Field Columbian Mus. Zool. Ser. (Chicago Nat. Hist. Mus. Fieldiana Zool.) Publication, 22: 319, fig. 9.

Type locality: hillside trickle between río Boquerón and río Pequeni, Chagres basin, Panama.

 

Cynodonichthys myersi (C. L. Hubbs, 1936)

Fishes of the Yucatan Peninsula. Carnegie Inst. Wash. Pub., 457: 210, pl. 2.

Type locality: 3 km south of Progreso, Yucatán, Mexico.

 

Cynodonichthys pacificus (Huber, 1992)

Review of Rivulus. Ecobiogeography - Relationships. Cybium Suppl., Société Française d'Ichtyologie Publ.: 351, fig. 67.

 

Type locality: Condoto, in forest along Condoto to Soledad road, 5 minutes driving from airport road, río San Juan basin, Colombia.

 

Cynodonichthys pivijay (Vermeulen, 2013) 

Four new species of Rivulus Poey, 1860, subgenus Cynodonichthys Meek, 1904 (Teleostei: Cyprinodontiformes: Rivulidae) from the Magdalena River Basin, Central Colombia, including notes on Rivulus (Cynodonichthys) elegans Steindachner, 1880. Killi-Data Series, 2013: 42-63.

 

 

Cynodonichthys pivijay "Col 2008-28"

 

 

 

 

Cynodonichthys ribesrubrum (Vermeulen, 2013) 

Four new species of Rivulus Poey, 1860, subgenus Cynodonichthys Meek, 1904 (Teleostei: Cyprinodontiformes: Rivulidae) from the Magdalena River Basin, Central Colombia, including notes on Rivulus (Cynodonichthys) elegans Steindachner, 1880. Killi-Data Series, 2013: 42-63.

 

Cynodonichthys rubripunctatus (W. A. Bussing, 1980)

Fishes of the Yucatan Peninsula. Carnegie Inst. Wash. Pub., 457: 210, pl. 2.

Type locality: 3 km south of Progreso, Yucatán, Mexico

 

Cynodonichthys siegfriedi (W. A. Bussing, 1980)

Status of the Cyprinodontid Fish Genus Rivulus in Costa Rica with Descriptions of new endemic Species. Brenesia, San Jose, 17: 340, figs. 2C, 5C, 6C.

Costa, W.J.E.M., 2003. Rivulidae (South American Annual Fishes). p. 526-548. In R.E. Reis, S.O. Kullander and C.J. Ferraris, Jr. (eds.) Checklist of the Freshwater Fishes of South and Central America. Porto Alegre: EDIPUCRS, Brasil.

 

Cynodonichthys sucubti (Breder, 1925)

New Loricariate, Characin, and Poeciliid Fishes from the Rio Chucunaque, Panama. Amer. Mus. Novitates, 180: 8, fig. 7.

Type locality: río Sucubti, in a small side stream, Darien province, Panama.

 

Cynodonichthys tenuis Meek, 1904 

The Freshwater Fishes of Mexico North of the Isthmus of Tehuantepec. Field Columbian Mus. Zool. Ser. (Chicago Nat. Hist. Mus. Fieldiana Zool.) Publication, 5: 101, fig. 27.

Type locality: El Hule (near Rio Papaloapan), Oaxaca (province), Mexico.

 

 

 

Cynodontichthys tenuis "Papaloapan"

 

Cynodonichthys uroflammeus (W. A. Bussing, 1980)

Status of the Cyprinodontid Fish Genus -Rivulus in Costa Rica with Descriptions of new endemic Species. Brenesia, San Jose, 17: 338, figs. 2B, 5B, 6B.

Type locality: San Vito landing field, tributary to río Coto Brus, Puntarenas Province, Costa Rica, 990 m altitude.

 

Cynodonichthys villwocki (Berkenkamp & Etzel, 1997)

Die Rivulus-Arten von Panama. 6. Zur Identität von Rivulus montium Hildebrand, 1938 mit der Beschreibung einer neuen Art, Rivulus villwocki spec. nov. D.K.G. (Deutsche Killifisch Gem.) J., 29 (6): 126, figs.

Type locality: near Salamanca, border of Chagres National Park, Colon province, Panama.

 

Cynodonichthys wassmanni (Berkenkamp & Etzel, 1999)

Die Rivulus-Arten von Panama. 7. Rivulus wassmanni spec. nov., eine neue Bachlingsart der Rivulus birkhahni-kuelpmanni-Gruppe aus der Provinz Bocas del Toro, Nordost Panama. Aquaristik aktuell, 5-6: 62, figs.

Type locality: 4 km west of Changuinola, Bocas del Toro province, Atlantic Panama.

 

Cynodonichthys weberi (Huber, 1992)

Review of Rivulus. Ecobiogeography - Relationships. Cybium Suppl., Société Française d'Ichtyologie Publ.: 455, pl. 16 (a-b) ; figs. 80, 82e.

Type locality: El Llano-Carti road, 12 km from IAH crossing, lago Bayano basin, (Pacific drainage) Panama.

 

Cynodonichthys xi (Vermeulen, 2013) 

Four new species of Rivulus Poey, 1860, subgenus Cynodonichthys Meek, 1904 (Teleostei: Cyprinodontiformes: Rivulidae) from the Magdalena River Basin, Central Colombia, including notes on Rivulus (Cynodonichthys) elegans Steindachner, 1880. Killi-Data Series, 2013: 42-63.