Aphyosemion (Mesoaphyosemion  25 Arten

Aphyosemion (Mesoaphyosemion) amoenum, Radda A.C. & Pürzl E. 1976

Der Aphyosemion cameronense-Komplex. DKG Journal 8 (11) 134-138.

Type locality: 3.5 km north-northeast of crossing Sondo-Dibang and Edéa-Yaoundé, west-central Cameroun.

Cameronense group 

Aphyosemion amoenum "Nyemtonye JVC 2013/19" (Kamerun 2013)

Fänger: Jaap Vlaming, Jaap Knol, Van Eck Joop und Heinz Ott


Aphyosemion (Mesoaphyosemion) aureum (A.C. Radda 1980)

Beschreibung zweier Aphyosemion Arten aus Süd-Gabun.

Aquaria 27 (11) : 201-205.)

Type locality: near Mouila, 47 km southwest of Koulamoutou, Du Chaillu massif, Ogooué-Lolo province, southern Gabon.

Aphyosemion aureum "Boudianguila GEB 1994/09" (Gabun 1994)

Fänger: Wolfgang Eberl & Thomas Blum

Aphyosemion (Mesoaphyosemion) cameronense, G.A. Boulenger 1903

(Descriptions of New Cyprinodont Fishes from Southern Cameroon. Ann. Mag. Nat. Hist. Series 7, 12 (70): 435-441, 1903.)

Bedeutung des Namens: nach dem Land Kamerun (Cameroon)

Type locality: Kribi river and Ja river (today Kienke and Dja), southern Cameroun 

Aphyosemion cameronense ABDEK 2012/474" (Gabun 2012)


Fänger: Agnèse Jean-François, Bogaerts Marc, Dening Cyrille, Eva Françoise und Kayoum Grégoire 

Aphyosemion cameronense "Mekaman JVC 2013/10" (Kamerun2013)


Fänger: Jaap Vlaming, Jaap Knol, Van Eck Joop und Heinz Ott 

Aphyosemion cameronense "Ngoyang" (Kamerun)

Aphyosemion cameronense "VAB 2016-11"  Fänger: Venstermans, Aerts & Bogaerts 

Aphyosemion cameronense "VAB 2016-21"  Fänger: Venstermans, Aerts & Bogaerts 

Aphyosemion obscurum "VAB 2016-31"  Fänger: Venstermans, Aerts & Bogaerts 


Aphyosemion (Mesoaphyosemion) citrineipinnis, J.H. Huber & A.C. Radda 1977

Cyprinodontiden Studien in Gabun IV Das du Chaillu Massif. Aquaria 24 : 89-110. 

Type locality: near Yéno, Mogambi, Ogoudou-Ogoulou sub-system of Ngounié, Gabon, ca. 450 m altitude. 

Aphyosemion citrineipinnis "Ekembele BSWG 1997-22"

 Fänger: Thomas Blum, P.Sewer, H.Weder R. Gluggenbuehl Juli / August 1997, Süd-Gabon


Aphyosemion (Mesoaphyosemion) coeleste (Huber & Radda 1977)

Cyprinodontiden-Studien in Gabun. IV. Das Du Chaillu-Massiv.

Aquaria (St Gallen), 24: 89-110, 10 figs., 5 tabs.

Type locality: near Mossango, 13 km northwest of Moanda, on way to Lastoursville, central Gabon, 400 m altitude.

Aphyosemion coeleste "GBG 1993/02" (Gabun 1993) Fänger: Friedrich Bitter & Wolfgang Grell.


Aphyosemion (Mesoaphyosemion) cryptum (Zee et al., 2018)

Aphyosemion (Mesoaphyosemion) cyanoflavum (Zee et al., 2018)

Aphyosemion (Mesoaphyosemion) mandoroense (Zee et al., 2018) 

Three new endemic Aphyosemion species (Cyprinodontiformes: Nothobranchiidae) from the Massif du Chaillu in the upper Louessé River system, Republic of the Congo

J.R.V.D. Zee, G. Walsh, V.N.B. Mikembi, M.N. Jonker, M.P. Alexandre and R. Sonnenberg

Zootaxa, 4369 (1): 63-92, 2018



Three new ‘Aphyosemion’ species are described from the upper Louessé River in the Massif du Chaillu, Republic of the Congo, based on a combination of DNA, habitat preference, male colour pattern, and morphological data. ‘Aphyosemion’ cyanoflavum, new species, is a member of the ‘A’. ogoense group. It differs from its congeners by a unique colour pattern and cephalic sensory system which contains a wide supra-orbital groove with large, densely pigmented anterior neuromasts and dark frontal neuromasts housed in one pit with one central anterior lobe. ‘Aphyosemion’ mandoroense, new species, and ‘A’. cryptum, new species, are members of the ‘A’. coeleste group. ‘Aphyosemion’ cryptum, new species, is in appearance very similar to ‘A’. coeleste, but lacks the typical post opercular metallic green blotch and is generally larger in body size. Initial DNA analyses demonstrate that ‘A’. cryptum, new species, is more closely related to ‘A’. mandoroense, new species, than to ‘A’. coeleste, despite similarity in appearance. ‘Aphyosemion’ cryptum, new species, and ‘A’. coeleste occur syntopic in several locations in a sub-catchment of the upper Louessé system, however differ in microhabitat preference. ‘Aphyosemion’ mandoroense, new species, differs by male body and fin colour pattern from all species of the ‘A’. coeleste group except ‘A’. citrineipinnis. From the latter, it can be distinguished by the absence of red pigmentation and a dark grey to black margin in the unpaired fins.


Aphyosemion cryptum (photo)  


Aphyosemion cyanoflavum (photo)    


Aphyosemion mandoroense (photo)  


Aphyosemion (Mesoaphyosemion) etsamense,  Sonnenberg &  Blum, 2004 

Aphyosemion (Mesoaphyosemion) etsamense (Cyprinodontiformes: Aplocheiloidei: Nothobranchiidae), a new Species from the Monts de Cristal, Northwestern Gabon.

Bonner Zoologische Beiträge. Band 53 (2004), Heft 1/2, Seiten 211-220.



Aphyosemion (Mesoaphyosemion) etsamense sp. nov., a species belonging to the A. (M) cameronense group, is described from the Monts de Cristal in northwestern Gabon. It is actually known from a small number of localities in the Monts de Cristal, which is also the westernmost known distribution area of the A. cameronense species group in Gabon. The new species differs from the geographically close populations of other species by the male colouration. The distinet status is also supported by mtDNA data. This is the second endemic nothobranchiid species described from the Monts de Cristal.

Bedeutung des Namens: nach der Stadt Etsam

Type locality: Etsam I, crossing the road N5 from Medoneu to Kougouleu, western slopes of the Monts de Cristal, Gabon.

Aphyosemion sp. aff. etsamense "GEM 2005/3


Fänger: Eberl und Mengila (Gabun 2005)
Aphyosemion etsamense "Etsam 1 GEB 1994/24"


Fänger: Wolfgang Eberl & Thomas Blum (Gabun 1994)


Aphyosemion (Mesoaphyosemion) haasi,

Radda  & Pürzl 1976

Der cameronense Komplex DKG Journal 8 (11); 140-141

Type locality: ca. 27 km northwest of Zomoko, Amvené tributary, north of Lalara, northern Gabon.

Aphyosemion (Mesoaphyosemion) halleri (Radda A.C. & Pürzl 1976)

Der Aphyosemion cameronense Komplex. DKG Journal 8 (11): 138-140.

Bedeutung des Namens: nach Ernst Haller

Type locality: Catholic Mission in Ambam, Cameroon.

Aphyosemion halleri " Ambam CSK 1995/21" (Kamerun 1995)

Fänger: Francisco Malumbres , Jorge de Torres, Bas Vlijm, Jos Keijzer & Geert-Jan van Huijgevoort


Aphyosemion halleri "Kekaman JVC 2013/10"

Fänger: Heinz Ott (Kamerun 2013)


Aphyosemion hanneloreae (Radda & Pürzl, 1985)

Zwei neue Formen der Gattung Aphyosemion aus Süd-Gabun. Aquaria (St Gallen), 32 (10): 157-160, fig.

Type locality: Malinga, Ngounié Province, near the border of Kongo, southern Gabon, 400 m altitud


Aphyosemion (Mesoaphyosemion) labarrei  (Poll 1951)

Notes sur les Cyprinodontidae de Musée du Congo belge. Premiére partie: Les Rivulini 

Revue de Zoologie et de Botanique Africaines 45 1-2): 168-170, map.

Bedeutung des Namens: nach Clement Labarre (Entdecker dieser Fische).

Type locality: creek of Ngufu river, near Kiavo, 20 km from Madimba (Inkisi), Zaïre.

Aphyosemion labarrei "AVD 2011 Kingembe" (Congo 2011)


X  𝐴𝑝ℎ𝑦𝑜𝑠𝑒𝑚𝑖𝑜𝑛 (Mesoaphyosemion) 𝑙𝑜𝑟𝑎𝑖, Stefano Valdesalici, Francisco Malumbres, 2022

A new killifish species from the Wele River system of Equatorial Guinea, Africa.

Published in aqua, International Journal of Ichthyology

https://www.aquapress-bleher.com/  (not valid ICZN)


Aphyosemion lorai "GEGUPO 2018 Fegayong"

Fänger: Adolfo González, Carlos Vizcaino, Francisco Portal, Heinz Ott


Aphyosemion (Mesoaphyosemion) mandoroense (Zee et al., 2018)

Three new endemic Aphyosemion species (Cyprinodontiformes: Nothobranchiidae) from the Massif du Chaillu in the upper Louessé River system, Republic of the Congo

J.R.V.D. Zee, G. Walsh, V.N.B. Mikembi, M.N. Jonker, M.P. Alexandre and R. Sonnenberg

Zootaxa, 4369 (1): 63-92, 2018 


Aphyosemion (Mesoaphyosemion) maculatum ,1977 A.C. Radda E. & Pürzl

Cyprinodontiden Studien in Gabun II. Nordgabon.

Aquaria 24: p 27-29, figure 7

Type locality: 33 km east Koumameyong, 20 km west Ovan, Koumameyong-Ovan National road No.4, northern Gabon.

Aphyosemion maculatum "ABDEK 2012/494" (Gabun 2012)

Fänger: Agnèse Jean-François, Bogaerts Marc, Dening Cyrille, Eva Françoise & Kayoum Grégoire

Maculatum group

Mesoaphyosemion losantosi Malumbres, Sonnenberg & Zee, 2022

Mesoaphyosemion montealenense Malumbres, Sonnenberg & Zee, 2022

Two new Mesoaphyosemion species from Equatorial Guinea (Maculatum group)


Mesoaphyosemion maculatum (Gabon)

Mesoaphyosemion losantosi (Equatorial Guinea)

Mesoaphyosemion montealenense (Equatorial Guinea)

Eschmeyer’s Catalog of Fishes

Valid as Aphyosemion maculatum, A. losantosi, A. montealenense


ABSTRACT Two new Mesoaphyosemion species and one new species of the ’Aphyosemion’ herzogi group are described from the inland of Equatorial Guinea. The results of mtDNA analyses of most of the known phenotypes of the genus Mesoaphyosemion and the ‘Aphyosemion’ herzogi species group, respectively, are presented. Both new Mesoaphyosemion species have dark blotches on the posterior flanks and resemble M. maculatum from Gabon, yet they are not closely related to that species. Although the two new species occur in very close proximity, DNA results suggest no close relationship. The ‘Aphyosemion’ herzogi species group has a similar distribution as the genus Mesoaphyosemion, but with its northern boundary in southern Cameroon. Based on mtDNA the new ‘Aphyosemion’ from the Mitemele drainage in south west Equatorial Guinea is basal species to the remaining species group. It is distinguished from the two described congeners, ‘A.’bochtleri and ‘A.’herzogi by a diagnostic combination of colouration characters. Unpaired fins and flanks have a green background and caudal peduncle is often yellow to golden with dark red irregular dark red bars. The genetic data indicate that the species group contains several additional, genetically and by colour pattern well separated, potentially undescribed species.




Aphyosemion (Mesoaphyosemion) mimbon, Huber 1977

Une chaine de deux Aphyosemion sympatriques dans les monts de Crystal, Gabon, avec description d'une

espece nouvelle; A.mimbon n.sp.

Revue Francaise d'Aquariologue 4 (1): 8-9.

Type locality: Akoga village, ca. 55 km (west) from Médouneu, flowing into the Mvé river in 150 m further down, Gabon.

Aphyosemion mimbon "BBS 1999/22" (Gabun 1999)

Fänger: F. Bitter, Blum & P. Sewer


Aphyosemion (Mesoaphyosemion) obscurum, 1924 E Ahl

Zur Systematik der altweltlichen Zahnkarpfen der Unterfamilie Fundulinae.

Zoologischer Anzeiger 60 (1-2): 55.

Type locality: Yaoundé, Cameroun.

Aphyosemion obscurum "ABDK 2010/426" (Kamerun 2010)

Fänger: Agnèse Jean-François, Bogaerts Marc, Dening Cyrille & Kayoum Grégoire

Aphyosemion aff. obscurum "VAB 2016-31"  Fänger: Venstermans, Aerts & Bogaerts 


Aphyosemion (Mesoaphyosemion) ocellatum, Huber & Radda 1977

Cyprinodontiden Studien in Gabon IV. Das du Chaillu Massiv. Aquaria Vol. 24: 99-100

Type locality: 6 km west of Mimongo, on way to Lébamba between Magagara and Lamadou, Migoto-Ogoulou system, Gabon.

Aphyosemion ocellatum "Nzenzele GBN 1988/17" (Gabun 1988)

Fänger: Friedrich Bitter & Roland Numrich.

Aphyosemion ocellatum Sika-Sika GJS 2000/05" (Gabun 2000)

Fänger: Robert Guggenbühl, Pascal Sewer & Mogens Juhl


Aphyosemion (Mesoaphyosemion) passaroi, Huber 1994

Aphyosemion passaroi, espèce inédite du Gabon sudoriental, au patron de coloration unique et description complémentaire de Aphyosemion decorsei (Pellegrin). Cyprinodontiformes, Aplocheilidae. Cyprinodontiformes, Aplocheilidae. Rev. Rev. Fr. Fr. Aquariol. Aquariol. v. v. 20 (núm. 3) for 1993 : 77-79. 20 (No. 3) for 1993: 77-79. Cyprinodontiden Studien in Gabon IV. Das du Chaillu Massiv. Aquaria Vol. 24: 99-100

Bedeutung des Namens: nach Guido Passaro

Type locality: 81.3 km east of Moukabou towards Koulamoutou, southeastern Gabon.

Aphyosemion passaroi "BB 2004/11" (Gabun 2004)  Fänger: Blum & Bitter


Aphyosemion (Mesoaphyosemion) punctatum, Radda & Pürzl 1977

Cyprinodontiden Studien in Gabon II. Nordgabun. Aquaria 24 : p 29-31, figure 8. Wildekampi group

Type locality: Makokou, near the Catholic Mission, northern Gabon.

Aphyosemion punctatum "Nbateka ABDK 2010/408"

Fänger: Agnèse, Bogaert, Dening, Kayoum (Kamerun 2010)


Aphyosemion (Mesoaphyosemion) raddai, Scheel 1975

Another new species of Aphyosemion from the Cameroons Journal of the American Killifish Association (9): p 259 263, map.

Bedeutung des Namens: nach Dr.A.C.Radda

Type locality: south of road junction Edéa-Eséka-Yaoundé, Cameroun

Aphyosemion raddai "ABL 2008/268" (Kamerun 2008)

Fänger: Agnèse Brummett & Lambert

Aphyosemion raddai "Matomb-Koktom JVC 2013/20"

Fänger: Jaap Vlaming, Jaap Knol, Van Eck Joop und Heinz Ott  (Kamerun 2013)


Aphyosemion (Mesoaphyosemion) wildekampi, Berkenkamp 1973

Beschreibung eines neuen Prachtkärpflings aus Aquarienfreund 2 (7): p 113-119, figures 3 & 4.

Bedeutung des Namens: nach Ruud Hans Wildekamp 

Type locality: vicinity of Diang, 40 km west of Bertoua, eastern Cameroun

Aphyosemion wildekampi  "VAB-2016-8"  Fänger: Venstermans, Aerts & Bogaerts


Aphyosemion wuendschi (Radda & Pürzl, 1985)

Zwei neue Formen der Gattung Aphyosemion aus Süd-Gabun. Aquaria (St Gallen), 32 (10): 157-160, fig.

Type locality: 50 km south of Mbigou, on the road to Malinga, Ngounié province, southern Gabon, ca. 600 m altitude.

Aphyosemion wuendschi "BSWG 1997/19" (Gabun 1997)

Fänger: T. Blum, P. Sewer, H. Weder R. Gluggenbuehl (Süden Gabon)